Education: M.S. (1970) Scripps Institute of Oceanography U.C., San Diego Ph.D. (1980) University of Rhode Island Research Interests: Research focuses on the physiological ecology of reef corals and on nutrient dynamics in tropical coastal systems. Past studies on corals have included: coral reproductive ecology and energetics; exchange of nitrogenous compouds between corals and their zooanthellae endosymbionts; nutrient effects on coral reefs.